A biologist who worked on cloning Dolly the sheep has died aged 58.
Nottingham University professor Keith Campbell famously created the sheep in 1996, together with fellow scientist Ian Wilmut.
He worked at Nottingham University from November 1999 until recently, where he held the position of Professor of Animal Development in the School of Biosciences.
Campbell first began researching animal cloning at the Roslin Institute near Edinburgh in 1991. The experiments led to the birth of Dolly the sheep.
Dolly was the first mammal to be cloned from an adult cell, and her creation sparked a debate about the ethics of cloning.
Dolly died in 2003.
